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In vivo ovulation, oocyte maturation and fertilisation in the bitch

Fontbonne, Alain (2008) In vivo ovulation, oocyte maturation and fertilisation in the bitch. PhD thesis Biologie de la reproduction, Ecole Nationale Vétérinaire d’Alfort, UMR 1198 Biologie de la Reproduction et du Développement, AgroParistech 2008AGPT0010 p.116.

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Abstract

Timing the day of ovulation as accurately as possible is considered by most authors as one of the

most important factor in order to determine when to inseminate bitches. This is especially important

when using frozen semen, due to the relatively short survival of frozen/thawed semen in the female

genital tract after artificial insemination.

In our study, we first tried to find the most accurate technique to determine the exact occurrence of

ovulation in the bitch. We hypothesized that ovarian ultrasound examination could be a reliable and

accurate method to determinate ovulation in bitches. Our aims were to try to detect ovulation by

ultrasound in different breeds and to evaluate the interests and the accuracy of such a technique,

compared with hormonal levels around ovulation. Our study confirmed that ovarian ultrasound was

an accurate technique for timing ovulation in the bitch. We also found that, although more accurate,

ovarian ultrasound examinations improved the accuracy of ovulation detection in only 15.3 % of

bitches compared with progesterone. Altogether, progesterone plasma concentrations appeared

fairly constant around ovulation, whatever the breed, and, as a whole, progesterone assays

appeared as a precise method to time ovulation.

Using ovarian ultrasound examinations as the reference technique to time ovulation in the bitch, we

then aimed to evaluate the precise kinetics of in vivo oocyte maturation in the bitch, as well as early

embryonic development. Furthermore, we wanted to check if, in vivo, spermatozoa were able to

penetrate oocytes still at immature stages. We found that the germinal vesicle (GV) stage was the

only one present until 44 hours after ovulation. The first metaphase II stage was observed at 54

hours. Various oocyte maturation stages were observed at the same time within each bitch.

Fertilisation occurred in most cases from 90 hours post-ovulation in mature oocytes (metaphase II).

In vivo penetration of immature oocytes by spermatozoa was extremely rare.

These fundamental results may lead to an improvement or a simplification in everyday veterinary

practice, especially for artificial insemination with frozen-thawed semen. Furthermore, a better

knowledge of the processes involved in in vivo oocyte maturation, fertilisation and early embryonic

development are important steps towards the improvement of reproductive biotechnologies in the

canine species.

Item Type:PhD Thesis (PhD)
PhD Supervisor:Chastant-Maillard, Sylvie
Date:14 February 2008
Board of examiners:Farstad, Wenche and Luvoni, Cecilia and Guérin, Pierre and Fiéni, Francis and Driancourt, Marc-Antoine and Guillot, Jacques and Chastant-Maillard, Sylvie
Ecole Doctorale:ED 435 AGRICULTURE, ALIMENTATION, BIOLOGIE, ENVIRONNEMENTS ET SANTE
Discipline:Biologie de la reproduction
Collection (Fonds):AgroParistech
Institution:AgroParistech
Department:Ecole Nationale Vétérinaire d’Alfort, UMR 1198 Biologie de la Reproduction et du Développement
Subjects:7. Life Sciences and Engineering
Uncontrolled Keywords:Dog, Chien, Ovulation, Oocyte, Fertilization, Fertilisation, Bitom, Insemination, Ultrasound, Ultrason, Ovary, Ovaire
ID Code:4418
Deposited By:Marina Briffaut
Deposited On:04 December 2008

Table of content

Introduction 11

In vivo oocyte maturation, ovulation and fertilisation in the bitch : state of the

art 13

Chapter 1. Determination of ovulation 25

Chapter 2. In vivo oocyte maturation and fertilisation 69

Chapter 3: Optimal timing for artificial insemination with frozen-thawed

semen 83

Discussion and perspectives 89

General conclusion 96

References 97

Annex 1 107

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